PhD Recruitment Process2022-03-23T12:28:59+00:00

The CRUK CoL Centre PhD Programme

Applications for the CRUK CoL Centre PhD programme starting in September 2022 are now closed!

This is a unique multidisciplinary and collaborative training opportunity for graduates with a background in biomedical or biological sciences, who are interested in carrying out research in cancer biotherapeutics.

Students accepted onto the programme will follow a four-year version of our PhD programme, benefitting from high quality multidisciplinary scientific & transferable skills training, and cohort-building & networking activities. In addition, you will develop a customised training plan together with your supervisor tailored to your background and project requirements.

We are looking for talented and motivated candidates, who are passionate about research and have a strong academic track record. You will be based in your primary supervisor’s research group, and you will have the opportunity to work across the Centre, for example when carrying out research placements or accessing the Centre’s infrastructure cores. These are full-time research positions fully funded for four years.

Eligibility criteria

Applicants must have obtained, or be about to obtain, a First or Upper Second Class UK Honours degree, or the equivalent qualifications gained outside the UK, in an area appropriate to the skills requirements of the Centre.

The funding for this fellowship covers students with Home tuition fee status only. For more information on Home tuition fee status please visit the UKCISA website. Please note that we will only be able to offer studentships to candidates that have Home tuition fee status or provide evidence that they can fund the international portion of the tuition fee from external sources (i.e. not self-funded).


Before beginning an application, you should read the information on this page and the “completing the application form” page. You should also find out about the projects being offered to start in September 2022 and their supervisors via the specific project pages below.

Key dates

  • Your application must be submitted by the deadline: 23 March 2022, noon
  • We should receive your references by 30 March 2022, noon. Your application may still be considered if references are not received by this deadline. However, no applicant will be invited to interview unless references have been received.
  • You will find out if you have been invited to interview for the programme by: 29 April 2022
  • Supervisors may contact you during the interview shortlisting period (late March to mid-April), to find out more about you and your interest in their project.
  • Information session for interviewees: 9 May 2022, noon to 1pm
  • Panel interviews will take place: 12 & 13 May 2022
  • Programme start date:  September 2022


During the interview, candidates will be interviewed by a panel of CRUK CoL Centre academics from across the Centre’s partners. Please note that not all projects will be funded and studentships will be awarded to the best applicants based on information submitted on the application form, references and performance during the interview.

PhD Projects available

Project 1: Investigating the roles of DNA Polymerase Epsilon in genome stability.

Primary supervisor: Roberto Bellelli, Queen Mary University of London. Co-supervisor(s): David Michod, UCL, Peter Cherepanov, Francis Crick Institute

Project 2: Nuclear and nuclear envelope remodelling during melanoma metastasis and immune control.

Primary supervisor: Jeremy Carlton, King’s College London. Co-supervisor(s): Victoria Sanz Moreno, Queen Mary University of London

Project 3: Post-transcriptional regulation of cytotoxic CD4+ T cell function in tumours.

Primary supervisor: Richard Jenner, UCL. Co-supervisor(s): Faraz Mardakheh, Queen Mary University of London

Project 4: Understanding myeloid heterogeneity in premetastatic niches following cytotoxic therapy.

Primary supervisor: Ioanna Keklikoglou, Queen Mary University of London. Co-supervisor(s): Sophie Acton UCL

Project 5: High-dimensional molecular analysis of tissue-resident macrophages and tier interactions with cancer.

Primary supervisor: Toby Lawrence, King’s College London. Co-supervisor(s): Chris Tape, UCL

Project 6: Untangling the mechanisms of presentation of unconventional epitopes to predict their immunogenicity and efficacy in anti-cancer immunotherapy.

Primary supervisor: Michele Mishto, King’s College London. Co-supervisor(s): Benny Chain, UCL

Project 7: Understanding mechanisms of glioblastoma invasion through manipulation of the tumour microenvironment.

Primary supervisor: Simona Parrinello, UCL. Co-supervisor(s): Sam Rodrigues, Francis Crick Institute, Ciaran Hill, UCL

Project 8:  Identification and characterization of tumour reactive TCRs for the advancement of cancer immunotherapies.

Primary supervisor: Sergio Quezada, UCL. Co-supervisor(s): Charlie Swanton, Francis Crick Institute

Project 9: Detection and targeting of early T and B cell dysfunction in pre invasive lung neoplasia.

Primary supervisor: James Reading, UCL. Co-supervisor(s): Timothy Tree, KCL

Project 10: Defining the Functions of EGFR Variants in Glioblastoma Heterogeneity.

Primary supervisor: Paulo Ribeiro, Queen Mary University of London. Co-supervisor(s): Alex Gould, Francis Crick Institute

Project 11: Evaluating targeting of the G2 catenation checkpoint as a novel tumour specific vulnerability.

Primary supervisor: Tanya Soliman, Queen Mary University of London. Co-supervisor(s): Miraz Rahman, King’s College London

Project 12: Imaging brain tumours to develop image guided focused ultrasound delivery of anti-PDL1.

Primary supervisor: Maya Thanou, King’s College London. Co-supervisor(s): Darren Hargrave, UCL, Elwira Szychot, Great Ormond Street Hospital, Antonios Pouliopoulos, King’s College London

Applications are now closed!